Ashley Greene
Credit: Jen Lowery / Splash News; Andy Fossum/Startracksphoto

Our favorite Twilight stars have come a long way since the first installment of the series back in 2008. Case in point: Ashley Greene, who originally auditioned for the Bella part but landed the Alice Cullen role, parlayed her popularity on and off the red carpet to become the star of Avon's Mark makeup and land the new DKNY spring campaign. With The Twilight Saga: Breaking Dawn Part 1 hitting theaters today, see how Greene and more stars have transformed over the past four years in the gallery!